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The Mokelumne River is a river in Northern California. The Upper Mokelumne River originates in the Sierra Nevada mountain range and flows into Camanche Reservoir in the Sierra foothills. The Lower Mokelumne River refers to the portion of the river below Camanche Dam. It crosses California's Central Valley, flowing westward through Lodi, California, and ultimately meeting its confluence with the San Joaquin River in the Sacramento-San Joaquin River Delta. The river crosses or forms the border of four California counties: Amador, Calaveras, San Joaquin, and Sacramento.